See Dipendra and Vaishali's paper on the evolution of multi-trait adaptations during the assembly of communities.
Basu, D. N., V. Bhaumik, and K. Kunte. 2023. The tempo and mode of character evolution in the assembly of mimetic communities. Proceedings of the National Academy of Sciences, USA, 120:e2203724120. Featured on the journal cover, in a PNAS Commenary, 'Butterfly mimicry rings run in circles', and in news articles on NCBS News (The Art of Subterfuge), Deccan Herald, and The Hindu.